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(2) Multidrug and toxic compound extrusion (MATE) transporters employ proton gradients to mediate flavonoid translocation into vacuoles or apoplastic spaces, exemplified by the Arabidopsis TT12 transporter functioning as a vacuolar flavonoid/H antiporter in seed coat cells accumulating proanthocyanidins (Sun et al., 2012)

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These findings underscore the complex bidirectional interactions between bone and brain, revealing potential therapeutic targets for cerebral ischemia and other neurological disorders (Figure 2)