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In vivo studies using two-photon fluorescence lifetime imaging and genetically encoded sensors (e.g., the laconic lactate biosensor) have confirmed that astrocytes generate lactate via glycolysis during neuronal activation, which is transported to neurons through monocarboxylate transporter MCT2

Metabolic reprogramming is characterized by mitochondrial biosynthesis dysfunction, increased glycolysis, and abnormal lipid and amino acid metabolism

Each analyte carries distinct physiological relevance: lactate as a marker of tissue hypoxia, creatinine for renal clearance, urea for nitrogen metabolism, and cortisol as an endocrine stress hormone [13,14]

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