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Additionally, the organic matrix plays a regulatory role in the biomineralization process of pearls, especially controlling the formation of CaCO 3 crystals (Ouyang, 2019), mainly manifested in organic matrix combination with specific crystal planes, leading to a reduced growth rate in this direction, and these slowly growing planes eventually dominate the morphology of crystals (Ma et al., 2012)

Importantly, it does not integrate into the host cells genome, eliminating mutagenic risks [141]

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Furthermore, the apoptosis-inducing effect and mechanism of FOXO4-DRI in a serum limitation-induced senescent fibroblast and organ culture models of keloids were examined