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It is well documented that ON bipolar cells hyperpolarize in response to glutamate, released in the dark, due the activation of mGluR6

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In general, the amino heterocycle strongly contributes to binding, in fact, the positively charged amino group forms a salt bridge with Glu467 (N-O distance: 2.7 for piperazine, 3.6 for piperidine), while the other main interactions are hydrophobic and involve Ile339, Phe396, Leu399 (Figure 4B)

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Am J Physiol 1979